The Last Fingerprint: How Markdown Training Shapes LLM Prose

Large language models produce em dashes at varying rates, and the observation that some models "overuse" them has become one of the most widely discussed markers of AI-generated text. Yet no mechanistic account of this pattern exists, and the parallel observation that LLMs default to markdown-formatted output has never been connected to it. We propose that the em dash is markdown leaking into prose -- the smallest surviving unit of the structural orientation that LLMs acquire from markdown-saturated training corpora. We present a five-step genealogy connecting training data composition, structural internalization, the dual-register status of the em dash, and post-training amplification. We test this with a two-condition suppression experiment across twelve models from five providers (Anthropic, OpenAI, Meta, Google, DeepSeek): when models are instructed to avoid markdown formatting, overt features (headers, bullets, bold) are eliminated or nearly eliminated, but em dashes persist -- except in Meta's Llama models, which produce none at all. Em dash frequency and suppression resistance vary from 0.0 per 1,000 words (Llama) to 9.1 (GPT-4.1 under suppression), functioning as a signature of the specific fine-tuning procedure applied. A three-condition suppression gradient shows that even explicit em dash prohibition fails to eliminate the artifact in some models, and a base-vs-instruct comparison confirms that the latent tendency exists pre-RLHF. These findings connect two previously isolated online discourses and reframe em dash frequency as a diagnostic of fine-tuning methodology rather than a stylistic defect.