What made useful knowledge cumulative was not discovery alone but the institutions that transmitted it. We provide the first exhaustive structural measurement of the network through which upper-tail human capital passed from master to student across a millennium. Using 470,000 mentor-student records from Wikidata (which integrates the Mathematics Genealogy Project and MacTutor Archive), and all 64 historical Fields Medalists as a fixed, ex ante tracer set, backward traversal yields a directed acyclic graph of 25.5 million paths reaching 57 generations. We document two institutional transitions. First, a 17th-century watershed concentrates lineage traffic on Leibniz: 47 of 64 lineages pass through him with a 10:1 downstream-to-upstream ratio, and seven independent attributes -- learned-society membership (a 46-fold rise per scholar), field, language, employer, institutional diversification, student production, and diffusion entropy -- re-organize coherently across the same window. This is the network signature of Mokyr's Republic of Letters, and it reframes the Newton-Leibniz priority dispute as a distinction between the possession and the transmission of upper-tail human capital: it is transmission that generates the spillovers on which growth depends. Second, 84% of lineages converge upstream on five 12th-13th-century Islamic and Byzantine scholars before terminating at an 11th-century boundary -- the ``Monastery Wall'' -- at which personal academic mentorship first becomes record-generating in Europe. Our claims are descriptive-structural, not causal. Because exhaustive traversal at this scale defeats standard tools, we also contribute a deterministic, algebraic graph-traversal instrument whose measurement bias we characterize in closed form, and report one emergent property of independent methodological interest.